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An application is typically a long-running service that is accessible over the network. Applications are the centre of a Juju deployment. Everything within the Juju ecosystem exists to facilitate them.
It’s easiest to think of the term “application” in Juju in the same way you would think of using it day-to-day. Middleware such as database servers (PostgreSQL, MySQL, Percona Cluster, etcd, …), message queues (RabbitMQ) and other utilities (Nagios, Prometheus, …) are all applications. The term has a specialist meaning within the Juju community, however. It is broader than the ordinary use of the term in computing.